1965-1968 Grundschule Hamburg "Am Lehmkuhlenweg"  
1968-1977 Gymnasium Athenaeum Stade  
1977 Abiturprüfungen  
  academic curriculum  
1977-1978 "studium generale" at Leibniz Kolleg, Tübingen  
1978-1979 Studies in chemistry at the University of Tübingen  
1979 Studies in biochemisty at the University of Tübingen  
1981 masters (diploma) in biochemistry  
1981/1982 eight-month stay in the Department of  Biochemistry at the University of Liverpool, England  
1984 diploma exam in biochemistry  
1984-1986 diplomawork at the Max-Planck-Institute for Developmental Biology in Tübingen (Department of Biochemistry; head: Prof. Dr. Uli Schwarz). Title: "Biochemical and immunological characterization of a glycoprotein from the extracellular matrix of the embryonic chick retina"  
1986-1989 PhD thesis at the Max-Planck-Institute for Developmental Biology. Title: "Layer formation in the tectum opticum of the chick embryo as exemplified by the separation of the retino-tectal and the tecto-bulbärer projection".  
February 1989 thesis defense in the Department of Biology/Zoology  
1989-1992 Postdoktoral fellow at the Department of Neurobiology, Stanford University School of Medicine, in the lab of Dr. U. J. McMahan. Title: "Cloning, characterization and expression of a chick agrin cDNA".  
1992-2000 head of an independent research lab at the Max-Planck-Institute for Brain Research in Frankfurt (Department of Neuroanatomy)  
1993-2000 teaching in the Department of Biology at the Technical University Darmstadt.  
1998 habilitation at the University of Darmstadt, "venia legendi" for Cell Biology and Neurobiology  
2000 - 2006 assistant professor at the Institute for Physiological Chemistry and Pathobiochemistry (medical faculty) at the University of Mainz  
2006 - 2007 Professor for Molecular Cell Biology at the University of Applied Sciences Lausitz (now called Brandenburg University of Applied Sciences)  
since 2007 Professor for the Physiology of adult neurogenesis in the Department of Physiology of the Medical School at the Ludwig-Maximilians-University Munich  
since 2007 Mentor in the Studienstiftung des Deutschen Volkes  
2016 Recipient of the "Felix-Jerusalem-Preis" of the Deutschen Gesellschaft für Muskelkranke e.V. ( for work on the development and pathology of the neuromuscular junction